Sub-aleurone layer

The endosperm cells are thin-walled and packed with amyloplasts containing compound starch granules. The endosperm is divided into two layers sub-aleurone and starchy endosperm. The two outermost cell layers (the sub-aleurone layer) are rich in protein and lipid they contain smaller amyloplasts and compound starch granules than the inner endosperm. The starch granules are mostly 3-9 pm in size. 

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