Genetic covariance additive

The generalized Fisher theorems derived in this section are statements about the space variation of the vectors of the relative and absolute space-specific rates of growth. These vectors have a simple natural (biological, chemical, physical) interpretation They express the capacity of a species of type u to fill out space in genetic language, they are space-specific fitness functions. In addition, the covariance matrix of the vector of the relative space-specific rates of growth, gap, [Eq. (25)] is a Riemannian metric tensor that enters the expression of a Fisher information metric [Eqs. (24) and (26)]. These results may serve as a basis for solving inverse problems for reaction transport systems. 

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