Ochyraea

Stech, M. and Frahm, J.-P. (2001) The systematic position of Ochyraea tatrensis (Hypnobartlettiaceae, Bryopsida) based on molecular data. Bryologist, 104 199-203. 

Vina, J. (1986) Ochyraea tatrensis gen. et spec, nov., a remarkable pleurocarpous moss from Czechoslovakia. Journal of Bryology, 14 261-267. 

Region R (position 137) has a shared substitution (G versus A) in 19 species. These included five of nine members of the LT clade (both species of Leskea, both species of Haplocladium, and Pseudoleskeopsis zippelii), four of seven manbers of the Lescuraea clade (LI) three of six species of Lescuraea (L mutabilis, L saxicola, L plicata), and Rigodiadelphus. Six additional species with this substitution Breidleria pratensis (012), Neodolichomitra yannanensis (04), Forsstroemia trichomitria (07), Ochyraea montana (OlO), Stereodon vaucheri and S. plicatulus (Oil)) were not related to the above groups or to each other (except the two Stereodon species). 

The type species of Ochyraea, O. tatrensis was found in a clade with O. smithii with high support (Nona 99% MB 100%). Relationship of these species was found previously by Stech and Frahm (2001), and some morphological characters, for example pseudoparaphylUa, are very similar in these two species. Interestingly, the opinion of R. Duell (Ochyra, personal communication) on the holotype of Ochyraea tatrensis was that this is just au aberraut form of Hygrohypnum smithii. A clade with Drepanium recurvatum and Tomentypnum nitens was found also by Tsubota et al. (2004). These two species were the only representatives of this clade in that analysis of approximately 600 species of mosses. 

Our previous analysis did not include Ochyraea (Hygrohypnum) mollis, thus this combination is introduced here as new Ochyraea mollis (Hedw.) Ignatov, comb. nov.-Hypnum molle Hedw., Species Muscorum Frondosorum TTi. 70 f. 7-10. 1801. 

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