Nucleotide functions energy charge

An important aspect of the function of photosynthetic complexes is their asymmetric arrangement in respect to the membrane and to the external and internal phases of the cellular compartments. This arrangement allows the catalysis of vectorial electron transfer and the performance of electrical work by promoting charge separation across the membrane dielectric barrier. It allows also in some cases the net translocation of protons across the membrane. These two processes are at the basis of the mechanism of energy conservation in photosynthesis coupled to the formation of ATP, which is added, in oxygenic photosynthesis, to the conservation of redox energy in the form of reduced pyridine nucleotide coenzymes. 

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