Are there alternative Fe II oxidation sites on ferritin molecules

7 Are there alternative Fe(II) oxidation sites on ferritin molecules  

A site is found in L ferritins from horse (HoLF) and human (HuLF) which is not present in the H ferritins, but is proposed to be an oxidation/nucleation site [4, 7, 8, 58], This site comprises glutamates 57, 60, 61 and 64 (H chain numbers) all of which lie on the inner siuface of the molecule [4, 7, 58]. Although faster than that of free iron, the initial rate of Fe(II) oxidation in HoLF or HuLF is about one-tenth that of HuHF and the L subunit is therefore unlikely to play a major role in Fe(II) oxidation in H, L copolymers. However in the heteropolymers, the combination of 

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